Research Areas

  • Fisheries
  • Aquaculture
  • Environmental Incidents
  • Fish Kills
  • Marine turtles
  • Marine mammal stranding
  • Benthic Ecology
  • Marine Productivity
  • Marine Reference Collection

 

The Fisheries and Aquaculture Research Programme conducts research in fisheries, aquaculture, and other aquatic biological resources.  A diversified programme in fisheries biology, ecology, conservation technology, genetics, age and growth and physiology emphasizes the integration of laboratory and field studies to develop scientifically sound approaches to the management of aquatic ecosystems.  Research is directed primarily towards the development of information and technology to increase understanding of aquatic ecosystems in Trinidad and Tobago and the Caribbean.  Technical assistance is provided to clients through Technical Advisory Services.

 




Projects

Fisheries

Fish Age and Growth Studies

  1. Age and growth studies of the wahoo, Acanthocybium solandri from the southern Caribbean (Trinidad and Tobago, Barbados, St. Vincent and St. Lucia).
  2. Age and growth studies of the Spanish mackerel, Scomberomorus brasiliensis from Trinidad in collaboration with the Fisheries Division, Ministry of Food Production, Land and Marine Affairs.
  3. Development of a Fish Otolith Reference Collection
  4. Photographic Atlas of Fish Otoliths from Trinidad and Tobago

The knowledge of age and growth is fundamental to fishery science.  The significance of determining age is that it allows fishery scientists to relate their observations to a time frame and estimate biological rates for various species.  Ages of individual fish are used to estimate growth rate, age at recruitment, maturity schedules, and age-specific fecundity for a specific species. In addition, the calculation of natural and fishing mortality rates and age-specific sex ratios also require age data.

This time frame may involve estimating the number of years a fish spends in a particular life stage or habitat, or determining the number of years that fishes are available for harvest.

While several methods exist (e.g. mark and recapture length frequency analysis) to determine the age and consequently the growth of individual fish and fish populations, the most robust method involves age determination from hard-parts of fish.  These hard-parts include scales, opercular bones, vertebrae, spines and otoliths (ear bones).  Ageing using hard-parts is the preferred route for age and growth determination because the resultant age estimates are more accurate and precise than those derived from length frequency analysis.

In the 1980s research on selected fish species was conducted at the IMA on an ad hoc basis.  However, in 1995, age and growth research was centralised with the formation of a Fish Age and Growth Laboratory which was established with assistance from a regional fisheries initiative, the CARICOM Fisheries Resource Assessment and Management Program (CRAMP).  Under this Program, age and growth information was provided for several commercial species by participating CARICOM countries.


From left to right: Annuli on whole sagitta of the crevalle jack, Caranx hippos, from Trinidad; Daily growth increments on lapillus of the Spanish mackerel, Scomberomorus brasiliensis from Trinidad; Annuli on sectioned sagitta of the Spanish mackerel, Scomberomorus brasiliensis from Trinidad and Annuli on sectioned sagitta of the lane snapper, Lutjanus synagris, from Belize.

 

Growth bands or rings (similar to growth rings on a tree), often called annuli can be seen on the hard-parts of fish.  These growth bands are often formed on an annual basis.

For most fish, otoliths or ear bones have been the most reliable indicators of age of all the hard parts.  Otoliths are formed by a process of biomineralization where calcium carbonate is precipitated on a protein matrix.  During periods of fast growth, the protein layers are more compact, giving  the growth zone an opaque appearance ,whilst during slower growth, these layers are more spaced, resulting in a translucent zone.  These two zones comprise an annulus. The deposition of annual growth rings is thought to be partly caused by seasonal changes in the environment and internal factors.

There are 3 pairs of fish otoliths - the sagittae, asterisci and lapilli.  The sagittae are more commonly used for determining ages formed on an annular basis whilst the lapillus is used to determine daily increments.  Ages can be determined from reading whole or sectioned otoliths.

 

Fisheries Co-Management Studies: An Investigation of Fisheries Resources, Resource Users and Fisheries Management by Communities to establish a Framework for Co-Management: Ortoire to Guayaguayare, Southeast Coast, Trinidad

This project was designed to identify and characterise the biological, technical and socio-economic attributes of this fisheries system and characterise and translate the views and knowledge of the fisheries system expressed by the fishers, fisheries scientists, managers and other resource users into a co-management framework which would allow stakeholders to address the prevailing problems in the fisheries and move to a more sustainable approach to its management.

The project was part of a larger regional initiative: Community–based Coastal Resource Management (CBCRM) Project funded by the International Development Research Centre of Canada (IDRC).

Collaborating agencies: International Development Research Centre of Canada (IDRC), International Ocean Institute of Costa Rica (IOI-Costa Rica), CARICOM Fisheries Unit of Belize (CFU), Laval University, Canada and the Institute of Marine Affairs.

Participating countries (CBCRM project): Trinidad and Tobago, Jamaica, Belize, Dominica, St Lucia, Venezuela, Panama, Mexico, Guatemala, Cuba and the Dominican Republic. Research methodology for the project included the use of face-to-face interviews guided by questionnaires to capture information on fishing operations, fisher households, and a perception and attitude survey on resource conditions and fisheries management issues.  Other research techniques included the use of key informants, focus group meetings, and participatory GIS involving cognitive mapping of fishing grounds, fish resources and boat migration patterns.

In addition, a video is being produced highlighting the fishing activities on the southeast coast of Trinidad and the development of the community-based co-management model used in the project.

(a) Pulling seine in Plaisance (b) Loading monofilament on boat for fishing trip (c) Setting seine boat
(d) Interviewing fisher for video on the fishing industry on the south east coast of Trinidad
(e-f) Co-management framework workshop conducted by the IMA with south east coast fishers in December 2006.

(g-h) Fish landing site at Guayaguayare (i) Young fisher-woman managing her father’s boat seine crew
(j) Women in Fishing Association (WIFA) general meeting (k) One of the fish species, Lutjanus synagris, lane snapper targeted by southeast coast fishers (l) One of the fish species, Scomberomorus cavalla, Kingfish, targeted by southeast coast fishers, Trinidad.

 

Fisheries and Participatory GIS

As part of the project, An Investigation of Fisheries Resources, Resource users and Fisheries Management by Communities to establish a Framework for Co-Management: Ortoire to Guayaguayare, Southeast Coast, Trinidad, cognitive mapping of fishing areas based on gear type as well as fish species was developed and incorporated in a GIS of the fishing system and other coastal activities from Ortoire to Guayaguayare.

Participatory GIS is a practice in which local communities share their knowledge and opinions to help generate maps to inform management and decision-making.

Cognitive mapping of fishpot fishing areas showing main fishing area, south east coast, Trinidad


Cognitive mapping by fishers of fishing grounds (by fish species), south east coast, Trinidad



Aquaculture

Aquaculture research at the IMA emphasizes the integration of laboratory and field studies to develop scientifically sound approaches to the management of aquatic organisms, production systems, ecosystems and aqua-businesses. The approach is to conduct research in aquaculture with the aim of satisfying the growing requirements of a needed industry which can assist in national food security, food production and the generation of sustainable aquaculture and related businesses.

Aquaculture research at the IMA started in 1984 with an overall goal of conducting field and laboratory trials and developing strategies for the commercial exploitation of targeted species. The species worked on initially were the apple snail Pomacea urceus, the cascadura Hoplosternum littorale, silver tilapia Oreochromis niloticus, red hybrid tilapia and the giant freshwater prawn Macrobrachium rosenbergii.

Fish Cages in Earthen Grow-out Ponds

Freshwater Prawns

Malaysian freshwater prawns Macrobrachium rosenbergii have been used in monoculture and polyculture trials at the IMA since 1985.  Research on prawn production from monoculture and polyculture with tilapia and cascadura, using locally produced feeds was conducted, and results including stocking densities, and survival and production rates, have been published in reports which are available at the IMA’s library. The prawn is an excellent prospect for polyculture and can be effectively used in polyculture systems with tilapia and cascadura.

Freshwater prawns Macrobrachium rosenbergii at different growth stages.

 

Tilapia

The IMA initially imported silver Oreochromis niloticus and red hybrid tilapia from Jamaica in 1985 and has further developed good brood stocks.  These have been donated to the Sugarcane Feeds Centre, Longdenville; the Bamboo Grove Fish Farm (Ministry of Food Production, Land and Marine Affairs); Orange Grove Fish Farm; the then Caroni (1975) Ltd. and also sold to private farmers.

A market survey conducted by the IMA showed that the red hybrid tilapia was a very popular fish with local consumers and compared very favourably with marine species in terms of colour, taste, texture, and price.

Both laboratory work and field studies are combined to give advisory services to the public, provide training, and produce seed stock for aquaculture development in Trinidad and Tobago.

Silver tilapia and red tilapia hybrid in dipnet at IMA Aquaculture facility

Aquaculture staff sampling fish in Tilapia Recirculating Aquaculture System (RAS)

Cascadura

Early aquaculture research at the IMA with the armoured catfish Hoplosternum littorale was geared towards food fish production systems, fry and fingerling production systems and larval rearing.  Work was completed in hatchery methodologies, production systems and marketing of aquaculture products for locally available species.  Researchers at the IMA have pioneered and developed methodologies for the commercial production of cascadura.  Investigations on the spawning and hatching of this species were conducted in an effort to improve nursery practices and to determine optimal stocking densities in production ponds. Cascadura can be stocked at relatively high densities and is suitable for polyculture with tilapia and freshwater prawns.  While natural stocks are being over-fished, there is an increasing demand for cascadura locally and also an export market.

Female and male cascadura in net seine at the IMA

 

Ornamental Fish

Trade in ornamental fish has been one of the most consistently expanding areas of aquaculture in Trinidad and Tobago.  Trinidad was once ranked third (behind Colombia and Brazil) in exports to the US from this region. Not all of Trinidad's exports are from actual local production and there is capturing of wild-caught fish. The practice of catching ornamental fish from our local rivers and streams for export is associated with environmental degradation, habitat loss and reduction in breeding areas.  This resource will probably be soon over fished, and farmers are encouraged to be involved in actual breeding and rearing of ornamentals.

Ornamental fish farm in Trinidad

Left: Ornamental fish in tanks at the IMA; Centre: Protozoan parasite Trichodina sp. (x 1,000); Right: Gill of stressed Angelfish Pterophyllum scalare (x 1,000).

The development of this industry will require a well-managed fish health programme.  Fish disease outbreaks are often complex, involving both infectious and non-infectious processes.  The research conducted by the IMA has enabled its specialists to provide much needed assistance to ornamental fish breeders as they seek to manage their businesses with greater efficiency in ways that do not damage the wider environment.

IMA Technician performing necroscopy on ornamental fish

 

Technical Advice

Technical advice to the public on all aspects of the fish production process and project cycle - from site evaluation, site selection, system design, species selection, pond construction, stocking, rearing, harvesting and marketing of aquaculture products is available through the Technical Advisory Services of the IMA.

 

Intensive Re-circulating Aquaculture Systems

Trinidad and Tobago is predominantly an oil-based economy with natural gas and petrochemical industries providing the bulk of Gross National Product (GNP).  Aquaculture has not been significantly supported or funded as an emerging industry, although there have been some successes in small-scaled fish farms and in ornamental fish breeding, farming and exports.  Traditional aquaculture systems which require available land, water and appropriate soil and topography have not stirred the imaginations of potential farmers and entrepreneurs or the local funding institutions for investment capital.   The cost of agricultural land is relatively high and successful enterprises are limited. However, increased demand for fish and fish products, coupled with declining stocks as well as the need for diversification in the local economy has seen a renewed interest in aquaculture as a source for fish and as a potential business.

 

Tank-based Tilapia Production System

The Institute of Marine Affairs (IMA) Fisheries and Aquaculture Research Programme (F&ARP) has introduced re-circulating aquaculture systems (RAS) to improve the efficiency and sustainability of aquaculture production systems.  One project based on this system is a collaborative effort with the Seafood Industry Development Company (SIDC) for the development of an intensive production system for growing hybrid red tilapias. There is no successful commercial aquaculture project for food fish in Trinidad and Tobago.  This project would serve to demonstrate the possibilities for successful commercial aquaculture and initiate further developments in commercial aquaculture.

Researchers wanted a flexible facility to raise tilapias from the fingerling to adult stage that met strict bio-security standards and which might also be used for other species. Production modelling and bio-programming for the project and final design and layout were prepared by staff of the F&ARP of the IMA.  Facility construction began in early 2009 and was completed in September 2009.  The facility is being run by IMA staff.

A covered shade-house of approximately 740 square metres and housing two separate production units comprising of six 30,000 litres tanks in each system, two 15 square-metre rooms – one for feed storage and the other a control room securing pumps, blowers, monitoring and alarm equipment -  and a shared covered area of 22 square metres between them was constructed.  Water is supplied by a 5 cm diameter PVC line from a well with capacity of 450 litres per minute use for supply and makeup water to all systems to satisfy bioplan requirements.  Recirculating systems include mechanical and biological filtration, aeration and on-demand drainage lines.  The wastewater treatment system designed for the facility includes solids filtration, water storage, fish and egg escapement prevention and management.  The system is equipped with water quality sensors, data-logging and alarm systems for effective management.  Culture systems are stocked with red hybrid genetically male tilapias (GMT) tilapias and silver GMT tilapia Oreochromis niloticus that were bred and raised in IMA hatchery systems.  Preliminary trials are ongoing with data collection on water quality, feeding, growth rates, fish health management, biofiltration, water budgeting and economics.  Initial results are encouraging.   Since May 2010, fish are being harvested on a weekly basis from the facility and are marketed by the SIDC.

 

Marine Re-Circulating Production System

Global marine fisheries have exhibited continuous declines in recent years and the Food and Agricultural Organization (FAO) has stated that 74% of the world's commercially fished species are depleted, over-fished or fully fished.  At the same time, due to the health benefits associated with a seafood-based diet, consumer demand for seafood has steadily increased.  It is very clear that in order to ease pressures on wild fisheries stocks and to satisfy the growing global consumption of seafood, marine species must be produced through aquaculture.  One such option is the use of mariculture for environmentally sustainable and economically feasible production systems.

Snappers and groupers are commercially viable fish species. Snappers and groupers belonging to the family Lutjanidae and Serranidae respectively have been cultured in marine cages, fibre glass tanks, and concrete tanks (indoors) or in ponds (outside) in the United States and Asia.  Locally, there has been little or no research on the potential of snappers and groupers as culture species.

The snapper species of importance are the southern red snapper Lutjanus purpureus, lane snapper L. synagris and the vermilion snapper Rhomboplites aurorubens while the grouper species most commonly targeted are the yellowmouth (Epinephelus flavolimbatus), and yellowedge grouper Mycteroperca interstitialis.  However, there are numerous species that may be cultured in Trinidad and Tobago which include: Lutjanus analis(mutton snapper), L. griseus (gray snapper), L. jocu (dog snapper), L. mahogani (mahogany snapper), L. synagris (lane snapper), L. purpureus (southern red snapper), Epinephelus adscensionis (rock hind), E. guttatus (red hind), E. fluvus (coney), E. itajara (jewfish), E. cruentatus (graysby), Mycteroperca venenosa (yellowfin grouper) and M. tigris (tiger grouper). Long-term and short-term trends in biomass for most of these stocks are unknown and there is no management plan for the fishing of grouper stocks in Trinidad and Tobago.  Anecdotal information suggests that stocks have declined and there is urgent need to put management measures in place.

Control Room for Marine Re-circulating System

The Marine Re-circulating Production System is part of the development of models for intensive, re-circulating aquaculture systems at the IMA using several species This project seeks to provide information on the survival and growth of several local snapper and grouper species in a marine re-circulating system as well as investigate the culture potential of other species.  It will also seek to promote intensive marine recirculating systems for the production of food fish and marine ornamentals as well as provide facilities for research in other marine organisms such as corals and sponges.

Activities will include collection of fish; acclimatization of fish into a marine re-circulating system; detection and management of pathogenic organisms by pre-examination of fish using histopathological methods; introduction of fish to commercially formulated feed and/or prepared rations; and determination of the feasibility as culture species in a marine re-circulating system. Preliminary trials have already been done and the Marine RAS is being scaled up for new trials beginning late, 2010.  Outputs expected are:

  1. IMA technical reports, short briefs and fact sheets to be published in the reference journal.
  2. A handbook on activities for extension work to aspiring marine aqua-culturists.
  3. Use of the in-lab model as a stepping stone for larger commercial tank grow-out and marine cages.

Publications

Publications

de Souza, G., and P. Gabbadon. 1992.  Cascadura Farming from Hatchery to Production.  Institute of Marine Affairs Aquaculture Series - 2.  Institute of Marine Affairs, Trinidad. 27p.


de Souza, G., and P. Gabbadon. 1989.  Feeding studies of cascadura fry Hoplosternum littorale Hancock 1828 under experimental conditions. IMA Research Report. Trinidad. 14p.


de Souza, G., and Gabbadon, P. 1991.  Fingerling production of the cascadura Hoplosternum littorale Hancock 1828.  IMA Research Report. Trinidad. 10p.


de Souza, G., P. Gabbadon and A. Titus. 1995.  Polyculture of giant fresh prawn (Macrobrachium rosenbergii) and cascadura (Hoplosternum littorale) in earthen ponds in Trinidad. Institute of Marine Affairs Technical Report. Trinidad. 9p.


de Souza, G., P. Gabbadon, and A. Titus. 1996.  A comparison of sinking and floating pelletized rations in the production of red hybrid tilapia.  IMA Technical Report. Trinidad. 8p.


de Souza, G., P.W. Gabbadon, and A. Titus. Polyculture Trials involving prawns Macrobrachium rosenbergii, armoured catfish Hoplosternum littorale, and red hybrid tilapia in earthen ponds in Trinidad.  47th Annual Gulf and Caribbean Fisheries Institute.


Gabbadon, P., and G. de Souza. 1990. Fish Farm Ponds – Design, Construction and Maintenance. 1990.  Institute of Marine Affairs Aquaculture Series - 1. Institute of Marine Affairs, Trinidad. 27p.


Gabbadon, P., and G. de Souza. 1989.  Preliminary Studies on Freshwater Prawn Macrobrachium rosenbergii Monoculture in Earthen Ponds in Trinidad. IMA Research Report. 23p.


Gabbadon, P. W., G. de Souza, and A. Titus. 2008.  Tilapia Farming: A Manual on Commercial Tilapia Production. Institute of Marine Affairs Aquaculture Series – 3 (2nd Edition) Institute of Marine Affairs, Trinidad.


Manwaring, G., G. de Souza, P. Gabbadon, and A. Titus.  1993.  Marketing the Red Hybrid Tilapia.  A Pilot Survey of Retail Fish Markets in Trinidad.  IMA Research Report.  Trinidad. 15p.


Gabbadon, P. and G. de Souza. 1985. The Freshwater Conch Pomacea urceus (Muller) as a Potential Culture Species for Trinidad and Tobago. In Proc. 38th Annual Gulf and Caribbean Fisheries Institute.


Gabbadon, P. W., and F. Chapman. 1996.  Use of the Lampricide 3-Trifluoromethyl-4-Nitrophenol (TFM) to Control Tadpoles in Warm-water Ornamental Fish Ponds. The Progressive Fish-Culturist 58: 23-28.


Kishore, R., and M. Antoine Malchan. (in Press) Local Fisheries Knowledge: A Case Study of Fishing Communities from Ortoire to Guayaguayare, Trinidad.  International Development Research Centre (IDRC) of Canada.


Kishore, R., C.M. Marshall, H. Ramsundar, G. de Souza, H. Haylock, and J. Finlay. 2006. Political organisation and socio-economics of fishing communities of Trinidad and Tobago, Belize and Grenada.  In: Coastal Resource Management in the Wider Caribbean: resilience, adaptation and community diversity.  Edited by Yvan Breton.   Kingston: Ian Randle Jamaica and International Development Research Centre, Ottawa, Canada.  265p.


Kishore, R., H. Ramsundar, and M. Clarke-Marshall. (in Press) Description of the fishery system of fishing communities from Ortoire to Guayaguayare, Trinidad. 253p. International Development Research Centre (IDRC) of Canada.


Kishore, R., H. Ramsundar, and M. Clarke-Marshall. (in Press) Fisheries Co-Management Framework: A Case Study of Fishing Communities from Ortoire to Guayaguayare, Trinidad.  International Development Research Centre (IDRC) of Canada.


Padmanaban, S.  (in Press) A GIS of the fisheries system and other coastal activities from Ortoire to Guayaguayare.   A case study of fishing communities from Ortoire to Guayaguayare, Trinidad. Prepared for International Development Research Centre (IDRC) of Canada.


Manickchand-Dass, S. 1987.  Reproduction, age and growth of the lane snapper, Lutjanus synagris (Linneaus) in Trinidad, West Indies.  Bulletin of Marine science, 40(1):22-28.


Manickchand-Heileman, S., and D.A.T. Phillip. 2000. Age and growth of the yellowedge grouper Epinephelus flavolimbatus and the yellowmouth grouper Mycteroperca interstitialis in Trinidad and Tobago. U.S. Fishery Bulletin 98, 290-298.


Manickchand-Heileman, S.C. and D.A.T. Phillip. 1996.  Reproduction, age and growth of the Caribbean red snapper (Lutjanus purpureus) in waters of Trinidad and Tobago.  In: F. Arregúin-Sanchez, J. L. Munro, M.C. Balgos and D. Pauly (eds.).  Biology, fisheries and culture of tropical groupers and snappers.  ICLARM Conf. Proc. 48:449p.


Manickchand-Heileman, S.C., and J.S. Kenny. 1990.  Reproduction, age and growth of the whitemouth croaker Micropogonias furnieri (Desmarest, 1823) in Trinidad waters.  Fishery Bulletin. U.S. 88:523-529.


Sturm, M., and G. de Souza. 1984.  Triportheus elongatus: A New record of a Characid Fish from Trinidad. In Copeia.  American Society of Ichthyologist and Herpetologists.


Sturm, M.de.L., and P. Salter. 1990. Age, growth and reproduction of the king mackerel Scomberomorus cavalla (Cuvier) in Trinidad waters.  Fishery Bulletin, v.88, n.2. 361-370.


Chin, X., and A. Richardson-Drakes. 2004.  Age and growth determination of the red hind Epinephelus guttatus from Grenada.  Document prepared for the CARICOM Fisheries Resources Assessment and Management Program (CFRAMP).  22p.


Kishore, R. 2005.  Age and Growth Determination of the lane snapper Lutjanus synagris from Belize and Jamaica. Document prepared for the CARICOM Fisheries Resources Assessment and Management Program (CFRAMP). 32p.


Kishore, R., X. Chin, and R. Ramsundar. 2003.  Potential Impacts of industrial development on coastal fisheries in the Gulf of Paria, Trinidad.  Proceedings of the 54th Annual Gulf and Caribbean Fisheries Institute. 13-27.


Kishore, R., and H. Ramsundar. 2008. Community-Based Fisheries Management: A Case Study of Fishing Communities from Ortoire to Guayaguayare, Trinidad. 59th Proceedings of the Gulf and Caribbean Fisheries Institute, Belize City: Belize. 59: 99 – 110.


Kishore, R., and F. Solomon. 2005.  Age and growth studies of Caranx hippos (Crevalle jack) from Trinidad using hard-parts.  56th Proceedings of the Gulf and Caribbean Fisheries Institute Conference.56: 227-239.

Partnerships

Partnerships

The IMA, through the Fisheries and Aquaculture Programme, has membership on the following Committees:

  • Management Committee of the Caribbean Fisheries Training and Development Institute (CFTDI)
  • Monitoring and Advisory Committee on the Fisheries of Trinidad and Tobago

Facilities

Early Aquaculture Facility at the IMA included:

  • A Wet Laboratory of approximately 500 square metres with thirty-two (32) 900 litre circular tanks and nine 0.02 ha outdoor earthen ponds
  • Water supplied by a flow-through well-water system from a well drilled on-site specifically for the project.
  • The well has the capacity to supply 120 gallons per minute of good quality water for the ponds and wet-laboratory.  It also supplements the IMA’s water supply.

Aquaculture Wet Laboratory at the IMA constructed in 1984/1985

 

Present-day Facility includes:

  • Aquaculture Laboratory
  • An Intensive Recirculating Aquaculture system comprising:
    • A shade-house
    • Ten 36,000 litre tanks
    • A feed storage area and control room
  • Benthic Laboratory
  • Fish Age and Growth Laboratory
  • Fisheries Laboratory
  • Histopathology Laboratory
  • Marine Productivity Laboratory
  • Marine Reference Collection Museum

Intense recirculating aquaculture system

Research Staff

Rosemarie Kishore

Senior Research Officer - Fisheries - This e-mail address is being protected from spambots. You need JavaScript enabled to view it

Farahnaz Solomon

Research Officer - Fisheries

Rhea King-Julien

Research Officer - Aquaculture- This e-mail address is being protected from spambots. You need JavaScript enabled to view it

Rachael Shoy

Research Officer- Fisheries Biology- This e-mail address is being protected from spambots. You need JavaScript enabled to view it